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The chapters of this book provide insights into the intricate mechanisms of nuclear import and export. To better understand these processes, one must first elucidate the organization of the physical gateways into the nucleus. this book begin with a detailed description of the nuclear pore structure and composition. The signal sequences that specify nuclear import and export of proteins are discussed next followed by eight chapters, each dedicated to a specific aspect of the nuclear import and export in plant and animal cells. Among these, special chapters are dedicated to nuclear import of Agrobacterium T-DNA during plant genetic transformation, nuclear import and export of animal viruses, and nuclear intake of foreign DNA. A chapter on research methods to study nuclear transport concludes the book. The result is a compact book which we hope the readers will find useful as a guide and a reference source for diverse aspects of nuclear import and export in plant and animal systems.